According to the information in the article, Kartik Sunagar and Yehu Moran

were the first scientists to concentrate their venom studies exclusively on ancient animal groups rather than on comparatively young animal groups.

were the first scientists to observe that venoms produced by ancient animal groups are more potent and complex than those produced by new animal groups.

were the first scientists to discover that the venoms of snakes and cone snails evolve through positive selection.

have proposed that negative selection is a more powerful and more prevalent evolutionary force than positive selection.

have proposed that the evolution of animal venom is better explained through the “two-speed” model than though the arms race analogy.